By declaring a tree you will gain:
Affliction - reduces the global cooldown of your Bane and Curse spells to 1 sec., increases your chance to hit with any spell by 3%, reduces spell pushback by 70% (any spell).So only Affliction currently has a unique bonus for specialising, but I expect that to be expanded to the other trees as development progresses. I'll try and get some Hand of Gul'dan action later in the day when the servers settle down (edit: looks like HoG is still tagged ingame as NYI, along with most of the new talents).
Demonology - increases your chance to hit with any spell by 3%, reduces spell pushback by 70% (any spell).
Destruction - increases your chance to hit with any spell by 3%, reduces spell pushback by 70% (any spell).
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